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Defining memory is about as difficult as defining time. In general terms, memory is a change to a system that alters the way that system works in the future. "A typical memory is really just a reactivation of connections between different parts of your brain that were active at some previous time," says neuroscientist Nikolay Kukushkin, coauthor of this paper. And all animals—along with many single-celled organisms—possess some sort of ability to learn from the past.
Like the sea slug. From an evolutionary perspective, you'd have a hard time drawing a straight line from a sea slug to a human. Yet they both have neurons, and sea slugs form something similar to memories. If you pinch a sea slug on its gills, it will retract them faster the next time your cruel little fingers come close. Researchers found synapse connections that strengthen when the sea slug learns to suck in its gills, and molecules that cause this change. Remarkably, human neurons have similar molecules.